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I am trying to discover the correct hyphenation and/or comma placement for the following sentence relating to honey bees:

When she hatches out of her egg, she is placed into a royal jelly filled hexagon cell called a queen’s cup.

I am inclined to punctuate as follows, but don't know if this is correct or why:

When she hatches out of her egg, she is placed into a royal-jelly-filled, hexagon cell called a queen’s cup.

If it is correct, can someone explain why? I believe the royal-jelly-filled part should be hyphenated because those words are related and would modify "cell." However, I am not as sure about placing the comma before "hexagon" or what the rule would be there, since it is an additional modifier of "cell."

I was previously pointed toward this post:

However, the examples in that post all seem to follow the two, or three word compound modifying the noun, such as brick-oven pizza or file-system-related software. In my case, I've got a fourth, unrelated modifier to deal with.

If my sentence were simply "When she hatches out of her egg, she is placed into a royal-jelly-filled cell called a queen’s cup" I would be satisfied with the hyphenation where I've placed it.

However, my sentence is actually:

When she hatches out of her egg, she is placed into a royal jelly filled hexagon cell called a queen’s cup.

Royal-jelly-filled is related. Hexagon is not. It's a separate adjective to describe the cell. Royal-jelly-filled-hexagon does not seem correct, so where then, does the hyphenation end? Does a comma belong after "filled"?

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  • Does this answer your question? Is it correct to hyphenate with compound premodifiers? If so, where is the hyphen placed? While J D OConal's answer (after Fowler?) would suggest << royal-jelly-filled hexagon cell >>, << file-system–related software >> (in the most upvoted answer) suggests << royal-jelly–filled hexagon cell >> (treating 'hexagon cell' as a compound). Commented May 8, 2023 at 10:11
  • Per @EdwinAshworth, the royal-jelly-filled part modifes hexagon cell, not cell alone. So the comma goes. Or reword it. Commented May 8, 2023 at 12:16
  • "Royal-jelly-filled is related." That's not how I read it; it made more sense to me for "royal" to modify "cell" and not "jelly" or "jelly-filled". However, I don't know anything about bees, so your interpretation certainly may be correct. Commented May 9, 2023 at 1:43
  • Isn't it a hexagonal cell and not the cell of a hexagon as your current wording implies?
    – tchrist
    Commented May 9, 2023 at 4:49
  • @tchrist 'Hexagon cell' is not unattested. Though not as well as 'Octagon Theatre'. Commented May 9, 2023 at 12:52

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