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Which of these sentences is correct?

"The meaning of these signs are well-understood."

"The meaning of these signs is well-understood."

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The 'meaning' is the subject of the sentence. It needs a verb in singular. So 'the meaning... is ...'

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If the signs all have the same meaning, then "The meaning of these signs is well-understood." makes sense, since meaning is singular.

If however you are talking about different meanings in a collection of signs, you should use: "The meanings of these signs are well-understood."

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