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According to Klaudia, birds disperse as an immediate response to flood, however they assemble back on the wetland, as the flooded areas elsewhere decline in extent.

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According to Klaudia, birds disperse as an immediate response to flood, however they assemble back on the wetland, as the flooded areas elsewhere decline in extent.

Yes Adrian, you need a comma after "however". However, you need to add a semicolon after "flood" as your sentence contains two independent clauses that relate to each other. Look at this:

According to Klaudia, birds disperse as an immediate response to flood; however, they assemble back on the wetland as the flooded areas elsewhere decline in extent.

The semicolon links "birds disperse as an immediate response to flood" to "however, they assemble back on the wetland as the flooded areas elsewhere decline in extent." It acts as a conjunction and works better for reader comprehension than two separate sentences would.

Notice I deleted the comma after "assemble back on the wetland" because "as the flooded areas decline in extent" is an adverbial phrase telling us when the birds assemble back on the mainland, and you usually don't use commas before adverbials.

Hope this cleared this up for you!

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