What is the proper way to punctuate a question that is posed within a statement like in the following example?
Associated with my interest in the patterns that structure aquatic ecosystems is the question of what factors control the distribution and cycling of resources in aquatic systems.
The second part of the sentence (after the what) is a question, but the sentence as a whole is a statement. Is the way I have it punctuated correct and what rules govern this sort of thing?