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What was going on with “quha”, “quhat” and the like in Scots and English?

From the Dictionar o the Scots Leid: Quha, Quhay, interrog. and rel. pron. Also: qwha, qha, qua, qwa, wha, vha, hua; qhaa; quhaw; quhai qwhay, whay, quay; quhae, whae; quhe, quhey, qwhey. ...