How would you capitalize a word in a book/article title when that word has an en dash in the middle of it? For instance, should "protein-protein interaction" be capitalized as "Protein-protein Interaction" or "Protein-Protein Interaction"?
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If each word is a word on its own, as in "Protein-Protein Interaction", yes. If the hyphen adds a prefix that doesn't stand on its own, like "Non-protein Elements", then I think not but I've seen it both ways.